Royal Society Publishing

Error message

Notice: Use of undefined constant all - assumed 'all' in include_once() (line 39 of /opt/sites/jnl-roybiogmem/settings.php).

John Alan Gulland. 16 September 1926-24 June 1990

R. J. H. Beverton, D. H. Cushing

This is a PDF-only article. The first page of the PDF of this article appears below.

PDF extract preview

Royal Society Login

Log in through your institution